The day unfolds at lunchtime, when Betty starts imaging things while she was eating a hot dog, when all of a sudden she sees Antonio Sabàto, Jr., then is interrupted by a image of Matt. Betty then tells Hilda that sooner or later she going to have to tell Matt about her final kiss with Henry.

It looks like MODE’s “Wedding Issue” have just been nominated for best cover issue at MOMAs. As Daniel is given good luck from Molly on getting the nomination, Wilhelmina gets wind that Claire is getting the credit, so Wilhelmina tries to use the info she got from Victoria as blackmail, demanding that Claire tender her resignation at the event or she’ll reveal Claire’s dark secret to Cal and the public about that tryst they had in Madison, South Dakota.

Daniel asks Betty to pay a visit to the features editor Penny Meadows, who seem to have her thoughts elsewhere, like complaining and wondering why all those piegons are on her window sill. Betty then gets a text from Matt to meet her downstairs. But just as Betty was finally going to say something to Matt, they witness someone falling from the 28th floor…Penny Meadows.

For Daniel, now that Penny has died, he agrees that Betty should get her job and says he’ll recommend it. While this is good news to the Suarez family, she tells them that she is cautious. The next day, a funeral for Penny is held, where everyone, especially Amanda, is shocked to learn that Penny has a twin sister, Mindy. Penny’s life is celebrated, sort of, with Betty giving the eulogy. However, Marc, who was offered the Vogue job but somehow turned it down, shows up to honor her with a big announcement that he will take over the feature editor job, shocking Betty in a big way. This causes Daniel and Wilhelmina to come up with a compromise: If they can come up with completing Penny’s research before the deadline, the winner will suceed Penny as feature editor.

As Betty prepares to track down Penny’s hardware at home, Matt stop by for lunch at City Park, only to reveal that he saw Betty kiss Henry at the same park bench they were sitting at as he lets his jealousy over Henry get the best of him. This prompts Betty to tell Hilda that he knows all about it.

At Penny’s home, Betty and Marc try to search for her notes when they are confronted by Mindy, who then takes her sister’s notes and throws it out of the window and orders the two to get out. This prompts Betty and Marc to come up with a plan by submitting resignations to Daniel and Wilhelmina as a way to buy time and decide on who can fill the position. Afterwards, Betty catches up with Matt to offer tickets, even though he was still bitter over his jealousy with Henry.

Meanwhile, Justin nervously waits to see if he got into performing arts school, hoping that year he’ll finally get accepted. However, when Ignacio gets the letter, he reads it and tells Justin that was not accepted, devastating the teen.

At the MOMAs, everyone is there, including Justin, who is wisked away by Marc, who gives him some advice about acceptance, which Justin takes to heart. Matt also shows up and has a talk with Cal, who offers him a job. Then there’s Daniel and Wilhelmina, who decided to flip a coin to come up with Penny’s replacement. Daniel later tells Betty that she has been promoted to features editor. While Betty feels excited over this, Marc does take it hard. Meanwhile, Cal comes to Wilhelmina to tell her that he and Claire knew about the pregnancy, and abortion, of their child, and it was none of her business.

Daniel is given good luck from Betty on getting the award when the wedding issue, with Molly on the cover as a bride, is chosen as the winner. Daniel then gives a speech without using his cue cards (which Marc took and gave to Wilhelmina) after he sees what he thought was Molly. As he went over to see if it was her, Betty told him that she didn’t see her. Daniel started to realize that the Molly he thought he saw in the audience is a ghost, as it turns out that she had died of cancer and that he learned the truth after her returned to the apartment.

Finally, more revealations awaited the parties involved. For Wilhelmina, she turns to a mysterious person to track down Connor Owens by any means necessary to retreive the money he stole from Meade Publications as a way to retaliate against the Meade-Hartley alliance, but as she walked into another room, her glass dropped as she sees someone in front of her. At the Meade mansion, Claire looked at a piece of paper that she would later burn in the fireplace, the paper being a adoption letter from South Dakota that reveals that she and Cal had a son and that he wasn’t aborted after all. Betty’s celebration as the new feature editor would turn sour when Matt stopped by to tell her its over between them, then stunned her with another surprise: he will now be working at Meade as her new boss. After Matt left, she gets a text from Daniel to come the apartment, where the two consoled each other.

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